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Table 1
The studied species diversity components and their descriptors. Indices based on presence-absence data are marked by pa .(1) Margalef (1958); (2) Heip (1974); (3) Berger and Parker (1970); (4) Shannon and Weaver (1949); (5) Simpson (1949); (6) Warwick and Clarke (1995); (7) Clarke and Warwick (2001).
| Component | Descriptor | Formula | Expected properties |
|---|---|---|---|
| Species richness | Species diversitypa | S=number of species | Standardize species richness by unit area |
| Margalef | 
|
Adjusted species richness by N (1) | |
| Evenness | Heip | 
|
Sensitive to rare species (2) |
| Berger parker | 1/d = N/Nmax | Sensitive to dominant species (3) | |
| Number of species + evenness | Shannon-Wiener | 
|
Sensitive to rare species (4) |
| Simpson diversity | 
|
Sensitive to dominant species (5) | |
| Species taxonomy | Taxonomic diversity | 
|
Extension of D including taxonomic relatedness (6) |
| Taxonomic distinctness | 
|
Form of limiting the influence of species dominance, reflecting pure taxonomic relatedness (6) | |
| Average taxonomic distinctnesspa | 
|
Equivalent of △ and △* in presence-absence data (7) | |
Variation in taxonomic distinctnesspaΛ+ where 
|
|
Evenness of the taxonomic level distribution in the (7) distinctness taxonomic tree |
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