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Table 1

The studied species diversity components and their descriptors. Indices based on presence-absence data are marked by pa .(1) Margalef (1958); (2) Heip (1974); (3) Berger and Parker (1970); (4) Shannon and Weaver (1949); (5) Simpson (1949); (6) Warwick and Clarke (1995); (7) Clarke and Warwick (2001).

Component Descriptor Formula Expected properties
Species richness Species diversitypa S=number of species Standardize species richness by unit area
  Margalef   Adjusted species richness by N (1)
Evenness Heip   Sensitive to rare species (2)
  Berger parker  1/d = N/Nmax Sensitive to dominant species (3)
Number of species + evenness Shannon-Wiener   Sensitive to rare species (4)
  Simpson diversity   Sensitive to dominant species (5)
Species taxonomy Taxonomic diversity   Extension of D including taxonomic relatedness (6)
  Taxonomic distinctness   Form of limiting the influence of species dominance, reflecting pure taxonomic relatedness (6)
  Average taxonomic distinctnesspa   Equivalent of △ and △* in presence-absence data (7)
Variation in taxonomic distinctnesspaΛ+ where    Evenness of the taxonomic level distribution in the (7) distinctness taxonomic tree

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